Jonas Lukošiūnas was born on October 7, 1919, to Veronika and Antanas Lukošiūnai in the Galiniai village, Užpaliai volost, Utena district. He was the eldest of six children. In 1932 he graduated from Užpaliai primary school after six forms, later lived and worked at his parents’ farm. He belonged to the organization “Pavasarininkai”, played in the Užpaliai wind band, and was a member of the Užpaliai Rifle Company. In 1941, Lukošiūnas became a member of the June Uprising. Jonas Lukošiūnas farm housed the rebel headquarters that was led by Juozas Namikas. In 1943, Lukošiūnas married Joana Gasiūnaitė with whom he had three children. In August of 1944, at the age of 25, he became a partisan. Lukošiūnas remained a partisan until July of 1946.